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F four other Wbl proteins (WblA, WblC, WhiD and WblE) occur in most actinomycetes (Figs 3 and 4), even though WblA and WhiD have developmental roles in S. coelicolor: WblA plays a key part in the transition of aerial hyphal initial branches to a sporulation-directed fate (wblA mutants have thin aerial hyphae often embedded in an extracellular matrix, with only occasional spore chains: FowlerGoldsw
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